Sorry to read about your failure. I started reading these posts a while back and honestly walked away from so much BS.
Something to consider. And others may correct me if wrong, but if the Spyder engine management could not bring the air fuel ratios into the correct range, whether with a stock exhaust or aftermarket exhaust, you would expect a fault code to show up and not clear itself.
I work with vacuum and pressure stuff almost daily.
Consider that if the fuel pump had created enough pull on the tank to develop a few inches of vacuum, and it is capable of a good amount, that is equal to a 1 psi of external pressure on the tank, the net surface area of the tank will crush since there will be hundreds of pounds net force crushing the tank from the outside based on surface area.
It was not heat, cold fuel, exhaust. Simply the vent system failed. As for the fuel cap, I just removed the cap on our 2014 RTs, the cap has no venting circuit at all. It is 100% a blocked cap and has no means to flow air at all.
You may want to see if mud dobbers built a nest in any of the hoses open ends, not the best place since there are gasoline fumes. But I have seen them build nests in fuel vents on other vehicles.
PK